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Freshwater Fishes of Iran Introduction Revised: Keys - active but still under construction. Bibliography note the papers listed in Published Accounts have updated bibliographies and contain references not found here. The families Adrianichthyidae Oryzias latipes , Percichthyidae Morone saxatilis , Centrarchidae Lepomis macrochirus and Micropterus salmoides , Mullidae Mullus barbatus , Scophthalmidae Psetta maxima and Pleuronectidae Platichthys flesus are deleted from this work as the species therein have not been confirmed from Iran see Coad and Abdoli b for some details.

Revised sections of this website appeared as review articles in the Iranian Journal of Ichthyology http: These families are no longer accessible here and will not be updated further. Some sections of this work are incomplete but have been posted as is.

They will be updated and queries resolved as time permits. Apart from files not yet complete, distribution maps, some figures and habitat photographs are the main items to be added. A question mark may appear in the text appended to items that need to be checked by me. Numerous queries have led me to post incomplete material.

This work has been carried out over a period of 45 years, starting in I arrived in Iran in January and, in that year, 7 articles were published strictly on Iranian fishes 3 on parasites, 1 on pesticides, 1 on fisheries, 1 describing the blind white fish and 1 a summary of the latter; 2 were in Farsi.

A generation later in , over articles on Iranian fishes appeared, along with hundreds of relevant works from neighbouring countries, works on the aquatic environment in Iran and works on taxonomy and systematics relevant to Iran. The study of fishes is now a very active field within Iran and the Middle East and much of the newer literature is easily available on-line see Bibliography. Accordingly, is the last year that this work was updated although some systematic and taxonomic studies may still be incorporated.

The Published Accounts see above were updated versions from content in this website. Acknowledgements A wide range of people in Iran, Canada and elsewhere have assisted me in this work over more than 40 years. Inevitably, I will have forgotten some names, which I regret.

Some people I never met formally, an example being the gentleman nattily dressed in suit by a stream near Kazerun who jumped fully-clothed into the water to help me catch fish. Numerous other Iranians have assisted my studies and this website is dedicated to them.

The staff at the Department of Biology, Shiraz then Pahlavi University helped me in numerous ways to collect fishes during a three-year tenure as an Associate Professor. Bahman Kholdebarin was Chairman of the Department for much of my time in Iran and it is only through his support that I was able to make the collections that enabled this work to be done. The Research Council of Pahlavi University funded field trips and is gratefully acknowledged for this support.

Collections were made with the help of drivers and assistants and their efforts over long periods in the field are gratefully acknowledged. Various other people assisted too and are mentioned below under the Pahlavi University name.

The staff in the CMN library searched out all the numerous and varied papers on fishes in Iran and neighbouring countries without which this synthesis would not be possible. One paper took six years to locate and arrived in the form of a microfilm from the Soviet Union.

I am particularly indebted to Victor Adomaitis who kindly volunteered for the unrewarding task of scanning hundreds of images and converting them to thumbnails and usable files. Mollie MacCormac carried on this task, making a wide variety of images available for the website. In particular, I should like to acknowledge the support and encouragement of the late Dr.

McAllister, Curator of Fishes, CMN over many years, in terms of training and education, both formal and informal, of financial and moral support, and in practical terms in the ways and means of collecting, cataloguing, identifying, and studying fishes, and of getting things done.

Co-authors are evident in the Bibliography and their added expertise made several studies possible. Various people and their organisations are mentioned below separately for their particular assistance; these are in alphabetical order. Asghar Abdoli collected numerous specimens including exotics and allowed me to incorporate these discoveries in several papers.

Behnke, Colorado State University, Fort Collins is gratefully acknowledged for his extensive loans of, and access to, collections he and associates made. These are listed more fully in the Materials and Methods. Bianco, University of Naples, allowed me free access to materials, including types, in his possession at the University of Naples and his hospitality is acknowledged. Petersburg are thanked especially for their hospitality, access to collections, data analyses and interpretations on Iranian fishes, as well as co-authorship.

Bond, Department of Fisheries and Wildlife, Oregon State University, Corvallis allowed extensive loans of fishes from Iran under his care and these materials are listed in the Material and Methods see Contents.

Staff at the Fish Section, British Museum Natural History now the Natural History Museum have loaned materials and hosted visits on numerous occasions; their help has been much appreciated for the extensive collections are a required study to understand the Iranian fauna. R Esmaeili, Shiraz University has contributed many items of information, DNA data, specimens and photographs, and has collaborated on a variety of studies on Iranian fishes.

Louis is acknowledged for his complementary studies on tooth-carps using molecular techniques. Kinzelbach kindly appointed me to the Advisory Board of the journal Zoology in the Middle East which has given me an interesting and valuable overview of studies in that region. Yazdan Keivany translated abstracts of his manuscript reports and first posted my bibliography of Iranian freshwater fishes on the internet - a stimulus to this work!

Bahram Kiabi, Gorgan University of Agricultural Sciences and Natural Resources is thanked for various items of information on fishes, translations and gifts of Farsi articles and many interesting fish specimens. His efforts at facilitating collegiality and his students have formed the core of modern university researchers on the fishes of Iran.

His published works are a model for students on fishes in that region. His enthusiastic cooperation in the field resulted in many interesting new specimens and his studies on distributions of fishes in Khuzestan have been very important for this web site.

The team at Ahvaz is acknowledged below individually and in teaching me Farsi names of fishes. Marammazi was head of that team and his hospitality and efforts to bring me to Iran are gratefully acknowledged.

Staff at the National Museum of Natural History Smithsonian Institution , Washington arranged loans of specimens and allowed access to the collections during several visits. Robin Ade, Stephane Ostrowksi, Ellis Pennington and David Currie sent specimens, photographs of fish and their habitats, drawings, and other data on fishes based on their field work in Afghanistan. Chris Shank sent, or arranged, the despatch of specimens and copies of documents as well as comments on fish names and other useful information.

Staff at the Fischsammlung, Naturhistorisches Museum Vienna have also loaned materials and hosted visits and their assistance has been essential to studies on Iranian fishes based on the collections of J. Jalal Valiallahi provided stimulating discussions on the limits and the content of the genus Barbus sensu lato in Iran while working at the CMN as well as a variety of photographs of these sometimes immense fish. Various people collected material for me or made gifts of material, sent specimens for identification, identified material, allowed access to collections under their care, made loans of material, provided other useful data and general information, and exchanged ideas.

These are listed below in alphabetical order with their affiliations at the time of their contribution sometimes only email addresses were known; and apologies if any titles are missing: Abd, Nature Iraq, Baghdad, H. Abedi, Savadkooh University, H. Alamdari, Organization of the Environment, Shiraz, A. Al-Attar, Basrah University, A. Al-Faisal, Basrah University, A. Al-Hakim, University of Nottingham, L. Ali, University of Basrah, S. Archibald, Yale University, Connecticut, Dr.

Armantrout, Portland, Oregon and family, Dr. Arratia, University of Kansas, Lawrence, S. Asadollah, Isfahan University of Technology, A. Balik, Ege University, Izmir, Prof. Balletto, Istituto di Zoologia, Genova, G. Beckman, Opelousas, Louisiana, Dr.

Berberian, Uinversity of Cambridge, Dr. Bertelsen, Zoologisk Museum, Copenhagen, Prof. Bohlen, Academy of Sciences, Libechov, Dr. Borkenhagen, Forschungsinstitut Senckenberg, Germany, Dr. Bornbusch, Duke University, Durham, Dr. Briggs, Watkinsville, Georgia, Dr. Cloutier, Department of the Environment, Tehran, Dr. Coffey, Pahlavi University, Shiraz, Dr. Smith Institute of Ichthyology, Grahamstown, Dr. Army Corps of Engineers, A.

Cross, University of Kansas, Lawrence, Dr. Dill, Davis, California, J. DouAboul, Kitchener, Ontario, Dr. Dugan, Penang, Malayasia, Dr. Ebrahimi, Lorestan University, Khorramabad, M.

Edmondson, Liverpool Museum, Dr. Ekingen, Veteriner Fakultesi, Elazig, O. Emamai, Shilat, Iran, Dr. Erk'akan, Hacettepe University, Ankara, Dr. Esmaeili, Shiraz University, E. Evans, Pahlavi University, Shiraz, K.

Fakhro, Directorate of Fisheries, Bahrein, R. Fazly, Fereydun Kenar, Mazandaran, R. Geerts, Swalmen, The Netherlands, Prof. Geldiay, Ege University, Izmir, Dr. George, Union College, Schenectady, Dr. Goren, Tel Aviv University, S.

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